Butterflies of Cuba
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    • Species List >
      • Papilionidae : Papilioninae >
        • Battus devilliers - De Villiers' Swallowtail
        • Battus polydamas - Polydamas Swallowtail
        • Heraclides andraemon - Bahaman Swallowtail
        • Heraclides androgeus - Androgeus Swallowtail
        • Heraclides caiguanabus - Poey's Swallowtail
        • Heraclides oviedo - Oviedo's Swallowtail
        • Heraclides oxynius - Cuban Black Swallowtail
        • Heraclides pelaus - Pelaus Swallowtail
        • Heraclides ponceana - Dusky Swallowtail
        • Neographium celadon - Cuban Kite Swallowtail
        • Papilio demoleus - Lime Swallowtail
        • Papilio polyxenes - Black Swallowtail
        • Parides gundlachianus - Gundlach's Swallowtail
        • Pterourus palamedes - Palamedes Swallowtail
        • Pterourus troilus - Spicebush Swallowtail
      • Hesperiidae : Eudaminae >
        • Aguna asander - Gold-spotted Aguna
        • Aguna claxon - Emerald Aguna
        • Autochton potrillo - Potrillo Skipper
        • Cecropterus dorantes - Dorantes Longtial
        • Chioides marmorosa - Cuban Longtail
        • Epargyreus zestos - Zestos Skipper
        • Phocides pigmalion - Mangrove Skipper
        • Polygonus leo - Hammock Skipper
        • Proteides maysi - May's Skipper
        • Proteides mercurius - Cuban Mercurial Skipper
        • Telegonus anausis - Caribbean Yellow-tipped Flasher
        • Telegonus cassander - Cuban Flasher
        • Telegonus cellus - Golden-banded Skipper
        • Telegonus habana - Frosty Flasher
        • Telegonus talus - Green Flasher
        • Telegonus xagua - Antillean Flasher
        • Urbanus proteus - Common Long-tailed Skipper
      • Hesperiidae : Hesperiinae >
        • Asbolis capucinus - Monk Skipper
        • Atalopedes mesogramma - Mesogramma Skipper
        • Calpodes ethlius - Canna Skipper
        • Carystoides mexicana - Mexican Ruby-eye
        • Choranthus radians - Radians Skipper
        • Cymaenes tripunctus - Three-spotted Skipper
        • Euphyes cornelius - Cornelius Skipper
        • Euphyes singularis - Singularis Skipper
        • Holguinia holguin - Holguin Skipper
        • Hylephila phyleus - Fiery Skipper
        • Lerodea eufala - Eufala Skipper
        • Nyctelius nyctelius - Violet-banded Skipper
        • Oarisma bruneri - Bruner's Skipperling
        • Oarisma nanus - Nanus Skipperling
        • Panoquina corrupta - Corrupt Skipper
        • Panoquina lucas - Purple-washed Skipper
        • Panoquina ocola - Ocola Skipper
        • Panoquina panoquinoides - Obscure Skipper
        • Parachoranthus magdalia - Magdalia Skipper
        • Perichares philetes - Caribbean Ruby-eye
        • Polites baracoa - Baracoa Skipper
        • Pyrrhocalles antiqua - Caribbean Skipper
        • Rhinthon cubana - Cuban Rhinthon
        • Saliana esperi - Perching Saliana
        • Synapte malitiosa - Caribbean Faceted Skipper
        • Wallengrenia misera - Cuban Broken-dash
      • Hesperiidae : Pyrginae >
        • Anastrus sempiternus - Common Anastrus
        • Burca braco - Braco Skipper
        • Burca concolor - Concolorous Skipper
        • Burca cubensis - Cuban Skipper
        • Burnsius crisia - Antillean Checkered Skipper
        • Burnsius oileus - Tropical Checkered Skipper
        • Chiomara gundlachi - Gundlach's Duskywing
        • Eantis munroei - Munroe's Sicklewing
        • Eantis papinianus - Cuban Sicklewing
        • Ephyriades brunnea - Florida Duskywing
        • Ephyriades zephodes - Zephodes Duskywing
        • Erynnis zarucco - Zarucco Skipper
        • Gesta gesta - Impostor Duskywing
        • Ouleus fridericus - Fridericus Spreadwing
      • Pieridae : Colladinae >
        • Abaeis nicippe - Sleepy Orange
        • Anteos clorinde - White-angled Sulphur
        • Anteos maerula - Yellow-angled Sulphur
        • Colias eurytheme - Orange Sulphur
        • Eurema amelia - Cuban Yellow
        • Eurema boisduvaliana - Boisduval's Yellow
        • Eurema daira - Barred Yellow
        • Eurema elathea - Banded Yellow
        • Eurema lucina - Smudged Yellow
        • Kricogonia cabrerai - Cuban Sulphur
        • Kricogonia lyside - Lyside Sulphur
        • Nathalis iole - Dainty Sulphur
        • Phoebis agarithe - Large Orange Sulphur
        • Phoebis argante - Apricot Sulphur
        • Phoebis avellaneda - Orange-washed Sulphur
        • Phoebis neleis - Neleis Sulphur
        • Phoebis orbis - Orbis Sulphur
        • Phoebis philea - Orange-barred Sulphur
        • Phoebis sennae - Cloudless Sulphur
        • Phoebis statira - Statira Sulphur
        • Pyrisitia chamberlaini - Chamberlain's Yellow
        • Pyrisitia dina - Bush Sulphur
        • Pyrisitia larae - Confusing Yellow
        • Pyrisitia lisa - Little Yellow
        • Pyrisitia messalina - Whitish Yellow
        • Pyrisitia nise - Mimosa Yellow
        • Pyrisitia proterpia - Tailed Orange
        • Pyrisitia venusta - Pale Yellow
        • Zerene cesonia - Southern Dogface
      • Pieridae : Dismorphiinae >
        • Dismorphia cubana - Cuban Mimic-White
      • Pieridae : Pierinae >
        • Ascia monuste - Great Southern White
        • Ganyra menciae - Cuban White
        • Glutophrissa drusilla - Florida White
        • Melete salacia - Black-striped White
        • Pontia protodice - Checkered White
      • Riodinidae >
        • Dianesia carteri - Caribbean Metalmark
      • Lycaenidae : Polyommatinae >
        • Brephidium exilis - Pygmy Blue
        • Cupido comyntas - Eastern Tailed-Blue
        • Cyclargus ammon - Nickerbean Blue
        • Cyclargus thomasi - Miami Blue
        • Hemiargus ceraunus - Ceraunus Blue
        • Leptotes cassius - Cassius Blue
        • Leptotes hedgesi - Hedges' Blue
        • Pseudochrysops bornoi - Antillean Blue
      • Lycaenidae : Theclinae >
        • Allosmaitia coelebs - Cuban Hairstreak
        • Chlorostrymon maesites - Amethyst Hairstreak
        • Chlorostrymon simaethis - Silver-banded Hairstreak
        • Electrostrymon angelia - Fulvous Hairstreak
        • Eumaeus atala - Atala Hairstreak
        • Ministrymon azia - Grey Ministreak
        • Nesiostrymon celida - Caribbean Hairstreak
        • Strymon acis - Bartram's Scrub-Hairstreak
        • Strymon bazochii - Lantana Scrub-Hairstreak
        • Strymon istapa - Mallow Scrub-Hairstreak
        • Strymon limenia - Limenia Scrub-Hairstreak
        • Strymon martialis - Martial Scrub-Hairstreak
        • Strymon toussainti - Toussaint's Scrub-Hairstreak
      • Nymphalidae : Apaturinae >
        • Asterocampa idyja - Dusky Emperor
        • Doxocopa laure - Silver Emperor
      • Nymphalidae : Biblidinae >
        • Dynamine postverta - Mexican Sailor
        • Dynamine serina - Caribbean Sailor
        • Eunica heraclitus - Cuban Purplewing
        • Eunica monima - Dingy Purplewing
        • Eunica tatila - Florida Purplewing
        • Hamadryas amphinome - Red Cracker
        • Hamadryas februa - Gray Cracker
        • Hamadryas feronia - Variable Cracker
        • Lucinia sida - Caribbean Banner
      • Nymphalidae : Charaxinae >
        • Anaea cubana - Cuban Leafwing
        • Archaeoprepona demophoon - Two-spotted Prepona
        • Cymatogramma echemus - Chestnut Leafwing
        • Hypna clytemnestra - Silver-studded Leafwing
        • Siderone galanthis - Red-striped Leafwing
      • Nymphalidae : Cyrestinae >
        • Marpesia chiron - Many-banded Daggerwing
        • Marpesia eleuchea - Caribbean Daggerwing
      • Nymphalidae : Danainae >
        • Anetia briarea - Many-spotted King
        • Anetia cubana - Cuban King
        • Anetia pantherata - Great King
        • Danaus eresimus - Soldier
        • Danaus gilippus - Queen
        • Danaus plexippus - Monarch
        • Greta cubana - Cuban Clearwing
        • Lycorea halia - Tiger Mimic Queen
      • Nymphalidae : Heliconiinae >
        • Agraulis vanillae - Gulf Fritillary
        • Dryas iulia - Flambeau
        • Eueides isabella - Isabella's Heliconian
        • Euptoieta claudia - Variegated Fritillary
        • Euptoieta hegesia - Mexican Fritillary
        • Heliconius charithonia - Zebra Heliconian
      • Nymphalidae : Libytheinae >
        • Libytheana carinenta - American Snout
        • Libytheana motya - Cuban Snout
        • Libytheana terena - Antillean Snout
      • Nymphalidae Limenitinae >
        • Adelpha iphicleola - Iphicleola Sister
        • Limenitis archippus - Viceroy
      • Nymphalidae : Nymphalinae >
        • Anartia chrysopelea - Cuban Peacock
        • Anartia jatrophae - White Peacock
        • Anthanassa frisia - Cuban Crescent
        • Antillea pelops - Antillean Crescent
        • Atlantea perezi - Cuban Checkerspot
        • Colobura dirce - Mosaic
        • Historis acheronta - Tailed Cecropian
        • Historis odius - Stinky Leafwing
        • Hypanartia paullus - Antillean Mapwing
        • Hypolimnas misippus - The Mimic
        • Junonia coenia - Common Buckeye
        • Junonia neildi - Mangrove Buckeye
        • Junonia zonalis - Tropical Buckeye
        • Phyciodes phaon - Phaon Crescent
        • Polygonia interrogationis - Question Mark
        • Siproeta stelenes - Malachite
        • Vanessa atalanta - Red Admiral
        • Vanessa cardui - Painted Lady
        • Vanessa virginiensis - American Lady
      • Nymphalidae : Satyrinae >
        • Calisto aquilum - Cuban Dark Calisto
        • Calisto bradleyi - Bradley's Calisto
        • Calisto brochei - Broche's Calisto
        • Calisto bruneri - Bruner's Calisto
        • Calisto disjunctus - Western Cuban Calisto
        • Calisto dissimulatum - Mimic Calisto
        • Calisto gundlachi - Gandlach's Calisto
        • Calisto herophile - Cuban Common Calisto
        • Calisto israeli - Israel's Calisto
        • Calisto lastrai - Lastra's Calisto
        • Calisto muripetens - Guamuahaya Calisto
        • Calisto occulta - Hidden Calisto
        • Calisto sharkeyae - Sharkey's Calisto
        • Calisto siguanensis - Sand Calisto
        • Calisto smintheus - Cuban Rusty Calisto
        • Calisto torrei - Torre's Calisto
    • Download Species List
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      • Papilionidae - Swallowtails
      • Hesperiidae - Skippers
      • Pieridae - Whites & Yellows
      • Riodinidae / Lycaenidae - Metalmarks, Hairstreaks & Blues
      • Nymphalidae - Nymphalids
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      • Stachytarpheta jamaicensis
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Another Milestone – Anolis lizards of Cuba

22/8/2019

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Picture
Anolis terueli Yellow-fanned Pallid Anole 28 Sept 2017 Camaguey © Tim Norriss
There are 64 species of Anolis lizards in Cuba that have been described to date but I’m sure that more will be described as more research is done on them. I think all are endemic to Cuba though some now occur in other places such as Florida or other islands as a result of deliberate or accidental introductions. Twenty years ago Lourdes Rodriguez Schettino produced the Iguanid Lizards of Cuba (1999). This book covered many topics including general taxonomy, morphology, ecology, genetics, parasitology, and biogeography. It covered 62 species but that included the iguana, curlytails and geckos, so many of the currently known Anolis are not featured. It featured colour illustrations of the males of just 50 species plus only four females and three immatures. The males, females and immatures in many species look completely different so in many ways it was nothing like our modern field guides. Whilst some are very restricted in range eg Anolis quadriocellifer, others like Anolis sagrei are widespread.
Anolis quadriocellifer Cuban Eyespot Anole 24 Jun 2015 Guanahacabibes © Tim Norriss
Anolis sagrei Cuban Brown Anole 20 Mar 2017 Topes de Collantes © Tim Norriss
I do find it surprising that nobody has since done a Field Guide to the reptiles of Cuba because it is desperately needed. The amphibians were covered in 2008 when Luis M. Díaz and Antonio Cadiz published Guía taxonómica  de los amphibious de Cuba. This was done through ABC Taxa and produced with financial support from the Directorate-General of Development Cooperation of Belgium. It was produced both in hardback and pdf versions and is freely downloadable from the ABC Taxa website. A possible reason for the time it’s taking to produce a similar guide to the reptiles is the desire to get more fieldwork done and gain more knowledge first. I hope this is not the case as in my opinion it is much better to publish sooner rather than later with the information that is known at that time. The increase in interest and knowledge amongst professionals and amateurs alike following publication is always worth it. There will always be new discoveries that add to or amend what was previously believed to be the case. That is fine and in the light of new knowledge there is no reason why a pdf edition cannot be quickly updated just as new revised editions of hard copy field guides are published at the moment.
Anoles can rapidly change their colour in response to external stimuli and I have watched this happen during interaction with another anole and during handling by a human. There are a few species that look superficially similar and it greatly helps the identification if the dewlap can be seen though of course this is not always possible. The four species below can all appear dark brown and so it is usually necessary to see the dewlap to identify it.
Anolis mestrei W Cuba © Raimundo Lopez-Silvero
Anolis homolechis male 27 Jun 2015 Soroa © Tim Norriss
Cuban Coast Anole Anolis jubar 2 Oct 2016 Guardalavaca © Tim Norriss
Anolis bremeri Cuban Variegated Anole 17 Jun 2015 Vinales © Tim Norriss
Picture
Anolis homolechis female (left), male (right) 17 June 2017 Vinales © Paul Bone
But it helps also to know the distribution of each species as that can also help to reduce the options. Modern digital photography has been a game-changer. I take lots of pictures of reptiles when we are out in Cuba and my knowledge of what we are seeing is gradually growing. For ones that I cannot identify I resort to the internet – and increasingly to Cuban contacts with way more experience than I have. For many of the scarcer species there are usually no photos anyway on the internet so I am very grateful to Nils Navarro for his recent help in identifying three species for me from our last two trips. The first is Anolis fugitivus - Green-headed Grass Anole. This is only known from the NSB mountains between Moa and Baracoa and has been confused with A. cupeyalensis in the past even by experts. It has been found on very few occasions but can be distinguished by having only 4 lines of raised scales across the dorsum rather than 6/7 in cupeyalensis.
Anolis fugitivus 11 Dec 2018 Moa © Tim Norriss
Anolis fugitivus 11 Dec 2018 Moa © Tim Norriss
Anoles can be divided into groups depending on what height above the ground they occur ie which habitat niche they have evolved to occupy. So Anolis fugitivus is considered a 'grass anole' and Anolis inexpectatus is one of the 'bush anoles'.
Anolis inexpectatus - Pineland Bush Anole 11 Dec 2018 Moa © Tim Norriss
Anolis inexpectatus - Pineland Bush Anole 10 Dec 2018 Moa © Tim Norriss
And the third species that Nils Navarro has kindly identified for me is Anolis pumilus - Cuban Spiny-plant Anole which we saw near Santa Clara in 2017 and again on Cayo Guillermo in June 2019 where it is right at the eastern end of its range.
Anolis pumilus 28 Mar 2017 Santa Clara © Tim Norriss
Anolis pumilus 15 Jun 2019 Cayo Guillermo © Tim Norriss
So what is the milestone? Well I have now just exceeded the halfway mark (33/64) of Cuban Anolis species with photographs on the Reptiles page of this website. There's a long way to go and getting photos of the other half will be a lot more difficult! Not all of the photos were taken by me and and I'm very grateful for any help I receive so if you have good quality photos of species not featured then I would very much welcome your contribution.
Picture
Anolis equestris 16 Jun 2018 Topes de Collantes © Tim Norriss
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